Penn’s homecoming victory over Brown on Saturday featured two elite wide receivers. One of them pulled in 17 catches for 170 yards and a touchdown. The other had 5 receptions for 82 yards and a touchdown.

Penn’s Homecoming game against Brown came down to the final minutes, as Connor Loftus kicked a 35-yard field goal to give the Quakers a 20-17 victory.

To beat Brown, which has one of the top defenses in the Ivy League, the Quakers will need to be more efficient inside the 20-yard line in Saturday’s showdown at Franklin Field.

No one has had a more up-close-and-personal view of the toast toss over the years than Donald Kelly. As head athletic groundskeeper, Kelly oversees the maintenance, painting and layout of all athletic playing surfaces at Penn.

Fitting with the season, the Quakers traveled to Yale on Saturday, only to find themselves immersed in a horror story.

Not every quarterback Penn has faced so far has been green, but the defense’s continued failures at making big stops is troubling, and even more so against those without much career playing time.

In a game marred by squandered opportunities and defensive struggles, Yale handed Penn football its first Ivy League loss of the season.

Though the Red and Blue have struggled with consistency so far this season, they still got the job done in their first two Ivy games. Yale has lost its last four games, but in order to avoid what coach Al Bagnoli called “a tale of two halves,” the Red and Blue will need to execute the full 60 minutes.

Andrew Holland and Eric Williams graduated from St. Ignatius High School in 2008 and 2011, respectively, and while playing for the Wildcats, they both led their squads to Ohio state championships.

Throughout the season, the Red and Blue have demonstrated the ability to mount comebacks late in games, and Saturday’s win fits that mold.

A contingent of researchers — including several from the Penn community — came together to make “Head Games,” a documentary about brain trauma in sports.

Penn’s inexplicable defensive collapse against Columbia’s last-ranked Ivy offense was alarming. The Lions moved the ball at will in the second half and racked up 475 total offensive yards in the game, almost 200 over its season average.

Trailing by three with 2:26 remaining on the game clock, Penn quarterback Billy Ragone led his team down the field, going 5-for-7 for 53 yards and finding senior running back Lyle Marsh in the end zone to give the Quakers a 24-20 win.

The threat of the pass is just what the doctor ordered against Columbia, since the Lions rank seventh in the Ancient Eight in pass defense efficiency.

Let’s just say this up front: The first four games of the Penn football schedule were rough.But here’s the good news: Columbia and Yale are up next.

After losing several key seniors following last season, defensive coordinator Ray Priore knew that some of his underclassmen would need to step in to fill their shoes. And just when duty called, Dan Wilk and Dan Davis emerged as stars of the Quakers’ young defense.

Freshmen Cameron Countryman and Dylan Muscat were fierce adversaries before they were teammates, playing on opposite sides of a heated rivalry between two public high schools in California. Muscat, a defensive back, often found himself matched up against Countryman, a receiver.