This marks the Quakers' first road loss of the season, while Harvard clinched at least a share of the Ivy title.
The two juniors made their mark all over the stat sheet as the team is still without two of its senior captains.
The game was tied at two following overtime, but a trio of missed penalties proved Penn's undoing.
The Quakers competed at four events following last spring's Ivy League Championship.
The Quakers won the conference last year following an excellent performance at the GEC Championships.
The Quakers head to Cambridge fresh off a stellar Homecoming showcase.
See how members of the DP's Sports Staff think the Quakers will fare in this season's final road test.
Brian Gill was named Coach of the Year, while five players were named to All-Ivy teams.
The Quakers are set to take on reigning Ivy champs, nationally ranked opponents, and century-old rivals.
The Quakers return Olympians, Junior National competitors, and Pan-American fencers.
As the top seed, the Quakers are hosting, and will Brown at Penn Park at 6:30 p.m.
The women's squad looks to best last year's success, as the men seek redemption.
This was the Quakers' fourth regatta of the fall, which also saw them travel to Boston for the Head of the Charles.
The freshman's 261 rushing yards set a Penn freshman record on Saturday.
Kevin Stefanski’s Cleveland Browns improve to 5-3 after bullying the Arizona Cardinals.
The Quakers finished second in the Open Eights race behind only Princeton.
Assistant coach Tyler Berger won gold for Team USA, while sophomore Adam Thomson represented Canada.
Seniors Maeve Stiles and Liv Morganti, and junior Dylan Throop led the Quakers with first-team recognition.
The recent Quaker graduate currently trains at the PRTC and represented Team USA.
Division III John Jay was no match for the barrage of new Quaker contributors.