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Nick Miller (#47) and teammates celebrate his interception returned for a touchdown to put his team up 21-7
Columbia coach Al Bagnoli
Senior Will Fischer-Colbrie struggles to find space as the Lions' defense swarmed the quarterback all game long.
Senior wide receiver Justin Watson catches the game-tying touchdown to force overtime
Junior defensive back Luke Bullock's dejected posture epitomizes the deflating loss, as the Quakers must now win the rest of their Ivy games to have any shot of claiming a third consecutive title.
Senior quarterback Will Fischer-Colbrie rushes for Penn's first touchdown against Columbia
Senior linebacker Louis Vecchio and the rest of Penn football's defense will look to rebound from a sloppy performance against Central Connecticut State University.
After subbing in for starting quarterback Will Fischer-Colbrie due to injury, sophomore quarterback Nick Robinson performed admirably in the second quarter, but was not able to help the Quakers overcome the CCSU defense in the second half.