Senior wide receiver Joe Holder (88) snags a touchdown pass from Billy Ragone in the third quarter. Holder had two catches for 30 yards, and Penn beat Yale 37-25.

Billy Ragone celebrates his fourth quarter touchdown run.

Wide receiver Ryan Calvert gets tackled by a Yale defender. Penn extended its unbeaten streak in the Ivy League to 18 games with a win over the Bulldogs Saturday.

Ryan Calvert (3) comes up just short on a pass from Billy Ragone. Calvert had a career day with 105 receiving yards on eight catches.

Penn defensive back Fred Craig (left) leaps to try to tackle Columbia running back Alec Fisher.

Penn wide receiver Ryan Calvert goes down two yards shy of the endzone.

Columbia quarterback Sean Brackett (10) scrambles away from Penn linebacker David Park (54)

Running back Jeff Jack gets upended by a Fordham defender before finding the goal line.

Junior running back Lyle Marsh (26) had a career night against Fordham, rushing for three touchdowns and averaging 5.1 yards per carry. Marsh split the running load with Jeff Jack as both filled in for Brandon Colavita, who sat out to avoid injury in the non-conference game. However, Marsh suffered an ankle injury in the fourth quarter.

Jason Schmucker returned an interception 48 yards to the Fordham 7-yard line to set up Penn for a score.

Sophomore center Chris Bush saw action on the junior varsity squad last year with fellow classmate and left tackle Jake Schwertner. The two have now moved up to the top of the depth chart for the Red and Blue’s all-important offensive line.

Junior safety Dan Ritt (13) fends off Dartmouth tight-end John Gallagher after intercepting a pass in the 3rd quarter to end a Dartmouth drive in Penn territory.

Quarterback Billy Ragone celebrates a game-winning touchdown with 17 seconds remaining at Dartmouth. Penn won 22-20.

Despite early-season struggles, junior quarterback Billy Ragone was in prime form Saturday night when he led two successful drives to end each half and put the Quakers in position to score.

Running back Brandon Colavita led Penn’s ground attack with 24 carries, topping the 100-yard mark for the first time this season. The Red and Blue picked up 206 total yards rushing in addition to 160 through the air.

Penn football fell to the Villanova Wildcats at Franklin Field (Villanova home game).

Penn football fell to the Villanova Wildcats at Franklin Field (Villanova home game).

Penn football fell to the Villanova Wildcats at Franklin Field (Villanova home game).

Junior quarterback Billy Ragone completed just 8 of 23 passes for 91 yards Saturday night in a blowout loss to previously winless Lafayette. Ragone threw two interceptions, both of which led to Leopards touchdowns, as the Quakers were overmatched on both sides of the ball.

Lafayette quarterback Andrew Shoop (back) passes over Penn senior Matt Hamscher.

Junior running back Brandon Colavita rushed 14 times for 67 yards, but Penn’s offense struggled through the air, completing just 17 of 40 passes.

Penn DB Anthony Marino (11) tackles Lafayette wide receiver Mitchell Bennett.

Third-string quarterback Andrew Holland saw some playing time against Lafayette as the game got away from the Quakers.

The tight ends pose for their photo at Penn football media day with tight ends coach Mark Fabish (center).

Starting quarterback Billy Ragone interviews with local media members during football media day.

The 2011 Penn football team poses for its annual team photo.

Penn football beats Princeton at Homecoming

Penn football beats Princeton at Homecoming

Penn football beats Princeton at Homecoming

Penn football is shut out at Brown, ending their Ivy streak.

Penn football is shut out at Brown, ending their Ivy streak.

Penn football is shut out at Brown, ending their Ivy streak.